A mysterious young woman, Hester Shaw, emerges as the only one who can stop a giant, predator city on wheels devouring everything in its path. Feral, and fiercely driven by the memory of her mother, Hester joins forces with Tom Natsworthy, an outcast from London, along with Anna Fang, a dangerous outlaw with a bounty on her head. In a post-apocalyptic world where cities ride on wheels and consume each other to survive, two people meet in London and try to stop a conspiracy.